Disaster / Destruction
When the Pillar of Heaven was shattered and the Sphaeran colonies were largely cut off from the galactic network by the Angelic blockade, the isolation proved too much for the burgeoning young settlements and their civilization collapsed; an event mirrored on harpiian colony worlds across the quadrant. After this, as the old research superprogram's experiments fell out of supervision, new creatures adapted to fill the niches opening up in the newly wild biosphere. This event is actually recorded in the planet's geology, denoted by the Sirpalean Line: a thin stratum of scattered microdebris from the Shattering, most visible in equatorial sediments.