The current glaciation period, called the Nehem glaciation, begins. During its first glacial cycle, most ulanids die out as their food sources disappear or their environment becomes too cold. However, two species manage to survive.
Those adapted to the brakish estuary and mangrove environments of southwestern Okaluan migrate further inland into the vast, hotspring fueled Ko forest swamps, adopting an amphibious, arboreal existence. These are the ancestors of the tua (Ulano feros).
Another species of ulanid adapted to the deep coral reefs at the bottom of the continental slope of eastern Okaluan escape into the deep sea vents along the Mid-Torvalen rift, quickly outcompeting other predators and adapting to the pressures of the deep. These are the ancestors of the abyssal ulan (Ulano tenebrae). Over time, their chromatophores have evolved to be phosphorescent in the lightless realms.
Excultids diverge into the Beccan and Labian genera. The Beccans spread northwards into Barendar. Labians emerge in Kemesh, some of which migrate to Okaluan and others into southern Kalmasa.